The England rugby coach Stuart Lancaster has called up uncapped forwards Mako Vunipola and Tom Youngs into his 32-man squad to play next month's matches against Fiji, Australia, South Africa and New Zealand.
The duo are two of four injury-enforced changes to the elite squad Lancaster originally named in July.
Saracens prop Vunipola, whose father played at two World Cups for Tonga, and Leicester hooker Youngs are covering for Alex Corbisiero and Rob Webber.
Harlequins winger Ugo Monye has also been promoted to fill the back-three berth left vacant when Ben Foden damaged ankle ligaments playing for Northampton.
James Haskell, who returned to the England team for the drawn third Test against South Africa in Port Elizabeth on the June tour, has been promoted from the Saxons to replace Tom Croft.