Captain James Horwill returns to the Wallabies starting line-up for the one-off test against Wales in Cardiff on Sunday morning as the teams meet in a rematch of their entertaining World Cup third-place playoff in October.
Horwill was named among the replacements for last weekend's 60-11 thrashing of the invitational Barbarians side at Twickenham with openside flanker David Pocock leading the team.
However, the Queensland Reds lock returns as skipper after a two-try, 30 minute cameo last weekend with Nathan Sharpe dropping to the replacements bench.
Centre Ben Tapuai and utility back Ben Lucas will both be optimistic of making their test debuts after being named alongside Sharpe on the bench.
Utility back James O'Connor continues at flyhalf after an impressive display against the Barbarians with inside centre Berrick Barnes recovering from a blow to his back to maintain the continuity in the backline.
Team: 15-Adam Ashley-Cooper, 14-Lachie Turner, 13-Rob Horne, 12-Berrick Barnes, 11-Digby Ioane, 10-James O'Connor, 9-Will Genia, 8-Ben McCalman, 7-David Pocock, 6-Scott Higginbotham, 5-Rob Simmons, 4-James Horwill, 3-Salesi Ma'afu, 2-Tatafu Polota Nau, 1-James Slipper.
Replacements: 16-Stephen Moore, 17-Ben Alexander, 18-Nathan Sharpe, 19-Radike Samo, 20-Ben Lucas, 21-Ben Tapuai, 22-Anthony Fainga'a