The Wallabies centre Rob Horne has withdrawn from the team to face Wales in Sunday's one-off rugby Test because of a tight hamstring and will be replaced by Anthony Fainga'a.
Horne failed to complete a training session at the Ospreys' base in Swansea.
Coach Robbie Deans promoted Queensland Reds centre Faingaa to the starting line-up and named Melbourne Rebels halfback Nick Phipps as the new bench man for the match at Cardiff's Millenium Stadium.
Phipps joins Ben Tapuai and Ben Lucas as Australia's backline reinforcements.
The pair of Bens are in line to become the third and fourth Test newcomers provided by the Queensland Reds this season, following Rod Davies (Wallaby No.851) and Beau Robinson (No.852) who debuted during the Test against Samoa in Sydney in July.