The New York Times has won five of this year's Pulitzer journalism awards.
The paper won prizes for investigative, breaking news and international reporting, as well as for feature photography and criticism.
The Las Vegas Sun won the prestigious public service award for reporting on the high death rate among construction workers on the Las Vegas strip.
Newspapers in the United States have been suffering from severe financial problems, with many publications closing, going bankrupt or making cuts.
They have also been struggling to compete with online publications.
Although these were allowed to compete for the first time this year, none of their 65 entries won prizes.
However, online content did play a role in a number of winning entries, the Pulitzer board said.
The New York Times, which itself is burdened by heavy debt, won the international reporting prize for its coverage of US involvement in Pakistan and Afghanistan.
Times journalist David Barstow was awarded the prize for investigative journalism for revealing how US networks used military commentators who had ties to the Pentagon or defence contractors and defended their policies.
The paper won the breaking news category for its work on a sex scandal that led to the resignation of former New York Governor Eliot Spitzer.
Named after newspaper publisher Joseph Pulitzer, who died in 1911, the accolades were first awarded in 1917.