Niger has turned a son of the late dictator Muammar Gaddafi over to Libyan authorities, the Tripoli government says, as a government-allied militia released pictures of him in captivity.
The government said on Thursday that al-Saadi Gaddafi, who fled across the Sahara desert to Niger during the 2011 uprising that saw rebels capture and kill his father, ending his four-decade dictatorship, was in custody in Libya.
The Tripoli Revolutionaries Brigade, a militia made up of former rebels, released pictures on Facebook of al-Saadi in a blue jumpsuit getting his head shaved, AFP reports.
The Libyan government said he would be held in accordance with "international standards regarding the treatment of prisoners."
Al-Saadi Gaddafi was best known as the head of Libya's football federation and a player who paid his way into Italy's top division.
Interpol had issued a "Red Notice" for him, for "allegedly misappropriating properties through force and armed intimidation when he headed the Libyan Football Federation."
Libya has repeatedly called for the extradition of al-Saadi from Niger, which had granted him asylum since September 2011 on "humanitarian" grounds, saying it had insufficient guarantees Libya's new rulers would give him a fair trial.
Three of Gaddafi's sons were killed in the 2011 uprising, including Mutassim, who was killed by rebels on the same day as his father.
Several key members of the Gaddafi clan have survived however, including Gaddafi's erstwhile heir apparent Seif al-Islam, who is wanted by the International Criminal Court but detained by a militia in Libya.
Former Libyan Olympic Committee chief Muhammad, and Hannibal, who made headlines with his scandal-packed European holidays, are believed to be in Algeria, as is the fallen tyrant's daughter Aisha.
About 30 senior regime officials are believed to have crossed into Niger at the same time as al-Saadi, but the authorities in Niamey have not said how many remain in the country.